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On the bolt thrower-mounted great knarloc, are the kroot aloud to fire their rifles whilst firing the bolt thrower?
Victory is achieved through mettle.
Glory is acheived through metal.
After a casual glance, it doesn't appear that the Riders can shoot their rifles, as they are effectively only a part of the Great Knarloc unit. Treat them as you would treat the crew pilot of a Rhino, for example :) You know he's in there driving, but he can't do much in game terms.